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Peter Arner is a leading figure in functional genomics, best known for his pivotal contributions to the FANTOM5 project, which revolutionized our understanding of gene regulation. His landmark 2017 paper, with over 360 citations, mapped transcription initiation events across the human and mouse genomes at single base-pair resolution using Cap Analysis of Gene Expression (CAGE) coupled with single-molecule sequencing. By analyzing nearly three thousand samples, Arner and his team created an unprecedented atlas of promoters and enhancers, revealing the dynamic landscape of transcriptional start sites across diverse cell types and conditions. This work has become a foundational resource for the genomics community, enabling researchers to decode how gene expression is controlled in health and disease. Beyond this flagship achievement, Arner’s research has consistently advanced our knowledge of transcription networks, making him a key architect of modern functional genomics. His meticulous, large-scale approaches continue to inspire new generations of scientists exploring the regulatory code of life.